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Leonato (Clark Gregg), the governor of Messina, is visited by his friend Don Pedro (Reed Diamond) who is returning from a victorious campaign against his rebellious brother Don John (Sean Maher). Accompanying Don Pedro are two of his officers: Benedick (Alexis Denisof) and Claudio (Fran Kranz). While in Messina, Claudio falls for Leonato's daughter Hero (Jillian Morgese), while Benedick verbally spars with Beatrice (Amy Acker), the governor's niece. The budding love between Claudio and Hero prompts Don Pedro to arrange with Leonato for a marriage.

In the days leading up to the ceremony, Don Pedro, with the help of Leonato, Claudio and Hero, attempts to sport with Benedick and Beatrice in an effort to trick the two into falling in love. Meanwhile, the villainous Don John, with the help of his allies: Conrade (Riki Lindhome) and Borachio (Spencer Treat Clark), plots against the happy couple, using his own form of trickery to try to destroy the marriage before it begins.

An elderly, arthritic former violinist named Marjorie, stares headlong into the decline of old age as her memories begin to fail her. Availing herself of a service that provides holographic recreations of deceased loved ones as their survivors would like them remembered, Marjorie spends her time with her daughter and son-in-law, and with an imperfect copy of her deceased husband, Walter, as he looked in his 30s and 40s.

Set in the near future, Marjorie Prime centers on Marjorie (Lois Smith), who spends time with the young likeness of her deceased husband Walter (Jon Hamm). This revitalized Walter is a sophisticated holographic projection that provides companionship while stimulating Marjorie’s memory—allowing her to explore their shared past as she lives with dementia.

The story revolves around Macbeth and Banquo, generals under the command of Duncan, the King of Scotland.